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  • competition from other species
  • damming of rivers
  • disease
  • drainage and filling of swamps and marshes
  • habitat destruction
  • hunting
  • hybridization with other species and subspecies
  • introduction of exotic species
  • leveling of forests for residential and industrial development
  • limited distribution
  • oil spills
  • overexploitation
  • overharvesting
  • poisoning
  • pollution
  • predation
  • strip mining
  • trapping
  • unregulated killing or illegal killing
  • water pollution

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